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Piper humistratum with blackish shiny leaves, Nouragues, CNRS field station, French Guyana
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Elatostema papillosum, a form with blackish leaves on forest floor, Putao, Kachin, Myanmar
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Elatostema papillosum, young shiny brown leaves later turning black in this blackish leaved form, Putao, Kachin, Myanmar
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Patrick Blanc and a black leaved form of Piper augustum, Calanoa, Leticia, Colombia, Nov. 2016
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Piper augustum, a form with almost black leaves, Calanoa, Letitia, Colombia
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Piper augustum, a form retaining black leaves at two meters obove forest floor, Calanoa, Letitia, Colombia
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Piper augustum, a form with almost black leaves and prominent nerves, Calanoa, Letitia, Colombia
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Piper augustum, close up of a black leaf, Calanoa, Letitia, Colombia
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Piper augustum, downward hanging maturing spadix and green undersurface of leaves, Calanoa, Letitia, Colombia
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Aspidistra babensis, population with blackish leaves, Ba Be NP, Vietnam, Nov. 2017
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Aspidistra babensis, population with its pachyphyllous blackish shiny leaves, Ba Be NP, Vietnam, Nov. 2017
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Aspidistra babensis with its pachyphyllous black shiny leaves, Ba Be NP, Vietnam, Nov. 2017
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Aspidistra babensis with black shiny leathery leaves, Ba Be NP, Vietnam, Nov. 2017
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Begonia cf. erythrogyna, black leaves and snow white dots, Poring, Kinabalu NP, Sabah, Borneo
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Begonia blancii, blackish form with slight blue iridescence, El Nido, Palawan, Philippines
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Begonia blancii, black leaved form, Palawan, Philippines
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Begonia blancii, dark brown spotted form, Palawan, Philippines
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Begonia blancii, light spotted form, Palawan, Philippines
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Begonia blancii, blackish, marbled and green individuals growing on the same rock, Palawan, El Nido, mai 2011
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Begonia blancii, mixed population of green, mottled, brown and blackish individuals, El Nido, Palawan, Philippines
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Begonia blancii, mottled and anthocyanic underleaf individuals, El Nido, Palawan, Philippines
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Homalomena humilis, population of blackish individuals, vertical earth slope, Maxwell Hill, Malaysia
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Homalomena humilis, blackish individuals, vertical earth slope, Maxwell Hill, Malaysia
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Amorphophallus sp. with blackish leaves, Hinboun, Laos
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Ipomoea sumatrana seedling with brown anthocyanic leaves in forest understory, Khao Yai NP, Thailand
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Ardisia cf. odontophylla, population with brwn glossy leaves, Selangor, Malaysia
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Ardisia primulifolia - Hong Kong
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Alocasia longiloba, dark purple leaf underleaf surface, Tioman, Malaysia
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Alocasia longiloba, dark purple underleaf surface with with whitish glands, Tioman, Malaysia
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Sonerila griffithii population with brown silver striped leaves on a mossy rock, seen from above, Gunung Ledang, Johore, Malaysia
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Sonerila griffithii, tuberous base and red anthocyanic leaf abaxial surface, Gunung Ledang, Johore, Malaysia
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Phyllagathis elliptica, red anthocyanic lower leaf surface, Kinabalu NP, 1600 m asl, Sabah, Borneo
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Amischotolype tenuis, big transparent epidermal cells, single chlorophyllous cells layer, mesophyll and lower epidermis with anthocyans in vacuoles giving brown leaf colour, Makokou, Gabon
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Columnea sanguinea, transparence,Tenorio, Costa Rica
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Columnea ericae, as an epiphyte with bright red leaf tips, Calanoa, Leticia, Colombia
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Columnea ericae, as an epiphyte just 3 m above forest floor, with bright red leaf tips and pale yellow flowers, Calanoa, Leticia, Colombia
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Columnea ericae with bright red leaf tips and pale yellow flowers, Calanoa, Leticia, Colombia
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Selaginella intermedia,close up of the brown form, the brown color originating in carotenoid pigments of rhodoxanthin group and not in anthocyans as in Angiosperms, Endau Rompin NP, Malaysia
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Selaginella intermedia, brown form, the brown color originating in carotenoid pigments of rhodoxanthin group and not in anthocyans contrary to Angiosperms, Endau Rompin NP, Malaysia
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Selaginella intermedia,close up of the brown form, the brown color originating in carotenoid pigments of rhodoxanthin group and not in anthocyans contrary to Angiosperms, Endau Rompin NP, Malaysia